Polystichum alfaroi
This species was locally abundant and was growing as an epiphyte in disturbed woods along the Rio Calderas above Boquete.

This species was locally abundant and was growing as an epiphyte in disturbed woods along the Rio Calderas above Boquete.
Wow! Is this a grammitid?
Yes, and weirdly enough, this clade is nested within Oreogrammitis and Radiogrammitis which look much more “typical”. See:
Sundue, Michael A., et al. “Global phylogeny and biogeography of grammitid ferns (Polypodiaceae).” Molecular phylogenetics and evolution 81 (2014): 195-206.
Cool! That’s a beautiful paper. I see that Oreogrammitis and Radiogrammitis are a mess…