Gymnocarpium has ca. 10 species and several hybrids. It occurs mainly in the temperate zones of the N Hemisphere (Asia, Europe, and North America) and subtropical mountains of Asia. The American G. dryopteris is a fertile allotetraploid species that arose from hybridization followed by polyploidization between G. appalachianum Pryer & Haufler and G. disjunctum (Ruprecht) Ching (Pryer & Haufler, Syst. Bot. 18: 150-172. 1993).
- habit – forest interior
- habit – forest interior
- habit – forest edge
- rhizome
- fiddlehead
- vascular bundles
- deltate lamina
- adaxial groove
- pinnae with swollen point of attachment
- free veins and sori
- sori












There is also a species from tropical Aisa, Gymnocarpium oyamense! I have photos from PNG and will post them soon.
Thank you, Michael. I have changed the text above. Looking forward to seeing your photos of G. oyamense.